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The 22 known Vp gene segments may be grouped into 17 different subfamilies, with each subfamily generally 36 ERIC LA1 ET AL. 9. Physical map of the mouse T cell receptor 0-chain locus. The Vp and Cp gene elements are indicated by solid boxes; the D and J gene segment clusters are indicated by lines. 3* is a pseudogene. The region of the 0-chain locus that is deleted in SJL mice is indicated by ASJL. , 1987). A Vp gene segment isolated by Singer et al. (1986)has been renamed as Val7 in agreement with the existing nomenclature.

Kronenberg, and E. h i , unpublished results). However, in contrast to the mouse 0-chain locus, it also appears that some V, gene segments have been duplicated over as much as 100 kb of DNA. , 1988). The repertoire of J, gene segments has been estimated at 50, although only 32 have yet been sequenced (Becker et a l . , 1985; J. 100 90 80 70 60 50 40 30 20 10 0 kb I FIG. 13. Physical map of the mouse a/6 locus. [For details on Ja sequences, see Arden et al. (1985), Winoto et al. (1985) (all numbers), Dembic et al.

The 21-hydroxylase B gene was found to be on the centromeric side and is about 300 kb distal to the DRa gene (Dunham et a l . , 1987) (Fig. 8a). , 1986). Long-rangephysical mapping has shown that TNFa and TNFP genes are located between the class I11 and class I regions. The distance between the C2 gene and TNFa is 34 ERIC LA1 ET A L . , 1988) (Fig. 8a). , 1987) (see Fig. 8a). The estimated total size of the human HLA locus is thus almost twice that of the mouse H-2locus (see above). The most significant difference in size was between the class I regions, where the homologous region in the mouse was about one-third that of man, despite the fact that genes homologous to Qa and Tla have not yet been found in humans.

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